| BEHAVIOR | CODE | DEFINITION |
| Agonism | ||
| Tufts-flick | TF | rapid back-and-forth movement of ear tufts |
| Frown | FR | lower eyebrows, furl brow, and turn down corners of mouth while staring |
| Cuff | CU | swift, superficial blow or scratch performed aggressively |
| Chase | CH | pursue partner, with one or both animals exhibiting aggression and/or submission (not play) |
| Fight | FI | grapple aggressively with partner(s), involving biting, clawing, and wrestling |
| Attack | AT | lunge at or pounce on partner aggressively; may or may not result in fight |
| Snap bite | SB | direct a single short, sharp bite at partner |
| Submit | SU | flatten ear tufts and/or facial grimace (partially open mouth with corners of mouth retracted, exposing lower and sometimes upper teeth) and/or slit eyes (eyelids half closed) |
| Continuous submit | CS | continuous submit; start scoring after 5 sec |
| Retreat | RE | starting from a stationary position, move at lest one body length away from another animal within 1 sec of the other animal establishing proximity (within 10 cm) |
| Play | ||
| Play | PL | two or more animals lunge, grapple, wrestle or chase for at lest 1 sec in absence of aggression or intense submission; play face may or may not be present |
| Solicit play | SP | direct play face toward, pounce on, or initiate grapple with partner, in absence of ongoing play with partner |
| Play face | PF | open mouth without retraction of the lips |
| Join play | JP | join ongoing play bout between two or more partners |
| End play | EP | discontinue all social play for _> 3 sec |
| Social play | SO | social interactions involving non-aggressive physical contact with other individuals; high activity |
| Infant-associated behaviors | ||
| Climb on | ON | climb onto any part of partner’s body so that all four limbs are on partner |
| Solicit climb on | SC | position body directly above infant and/or pull infant onto body; may or may not result in infant climbing onto partner’s body |
| Climb off | OF | voluntary climb off partner’s body after having all four limbs on partner |
| Push off/reject | PO | prevent juvenile from climbing onto body, or rub or otherwise force juvenile off body |
| Nurse | NU | have mouth on female’s nipple for _> 1 sec |
| End nursing | EN | discontinue nursing posture |
| Other social behaviors | ||
| Sniff/nuzzle | SN | orient face against or toward partner, excluding anogenital region |
| Anogenital inspect | AI | orient face against or toward anogenital region of partner, or use hands or mouth to investigate anogenital region of partner; includes anogenital groom |
| Groom | GR | use hands and/or mouth to pick through fur and/or mouth of partner, excluding anogenital region |
| Sexual solicit | SS | stare at partner with ear tufts flattened and eyes slit |
| Mount | MO | climb on partner’s back from behind and grip partner around waist and legs; may be accompanied by pelvic thrusting |
| Initiate huddle | IH | establish passive, torso-torso body contact with partner, with both animals remaining stationary and in passive contact for at least 3 sec |
| Leave huddle | LH | terminate huddle after at least 3 sec of passive, torso-torso body contact during which both partners remained stationary |
| Object steal | OS | take any non-food object from hands or mouth of partner |
| Attempt object steal | AO | attempt but fail to take non-food object from hands or mouth of partner |
| Food-associated behaviors | ||
| Food steal | ST | take any food from hands or mouth of partner |
| Attempt food steal | AF | attempt but fail to take food from hands or mouth of partner |
| Share food | SH | eat from a food source from which partner is simultaneously eating or occupying without removing any food from partner’s mouth or hands |
| New food | NF | eat from a food source which no other animal is currently holding, eating from, or occupying |
| Individual behaviors | ||
| Bristle strut | BS | arching posture and/or strut locomotion and/or general Piloerection |
| Scentmark | SM | rub or drag anogenital, suprapubic, or sternal region along substrate, object, or partner |
| Genital present | GP | raise tail to expose genitals |
| Object manipulation | OM | sniff, bite, chew, gouge, handle, pounce on, grapple with, or otherwise manipulate inanimate object, excluding food items and water bottle, for at least 1 sec |